Physiology

Cardiac Physiology

Cardiac cycle, action potentials, output regulation, and hemodynamics

The Cardiac Cycle

The cardiac cycle includes systole (contraction) and diastole (relaxation) coordinated by electrical signals.

PhaseValvesKey Events
Isovolumetric contractionAll closedPressure rises, no volume change; follows QRS on ECG
Rapid ejectionAortic/pulmonic open70% of SV ejected; peak systolic pressure
Isovolumetric relaxationAll closedPressure falls, no volume change; follows T wave
Rapid fillingMitral/tricuspid open80% of ventricular filling (passive); S3 sound here if pathologic
Atrial kickMitral/tricuspid openFinal 20% filling; S4 sound here if pathologic; lost in A-fib
⭐ Heart SoundsS1 = mitral/tricuspid closure (systole onset). S2 = aortic/pulmonic closure (systole end). S3 = rapid filling (volume overload, HFrEF, normal in young). S4 = atrial kick against stiff ventricle (hypertension, LVH, AS).

Cardiac Action Potentials

PhaseIon FlowSignificance
Phase 0 — Rapid depolarizationNa+ in (fast channels)Absent in SA/AV nodes; blocked by Class I antiarrhythmics
Phase 1 — Early repolarizationK+ out (Ito)Notch on AP; prominent in Brugada syndrome
Phase 2 — PlateauCa2+ in, K+ out (balance)Unique to cardiac muscle; sustains contraction; blocked by Class IV (CCBs)
Phase 3 — Rapid repolarizationK+ out (IKr, IKs)Blocked by Class III antiarrhythmics (amiodarone, sotalol); prolongs QT
Phase 4 — Resting potentialK+ leak; pacemaker If (funny current)SA node spontaneous depolarization; blocked by ivabradine
🩹 SA vs Ventricular APSA/AV node AP: no Phase 0 fast Na+; relies on slow Ca2+ channels (Phase 0). Slow conduction. Blocked by Class IV (CCBs) and Class II (beta-blockers). Ventricular AP: fast Na+ dependent; Class I antiarrhythmics work here.

Cardiac Output and Determinants

CO = HR x SV (normal ~5 L/min at rest)

DeterminantDefinitionIncreases WithDecreases With
Heart rateBeats per minuteSympathetic, catecholamines, hyperthyroidismParasympathetic (vagal), beta-blockers, hypothyroidism
PreloadVentricular EDV (stretch)Increased venous return, fluid bolusDiuretics, venodilatation (nitrates)
AfterloadResistance ventricle pumps againstHypertension, aortic stenosis, vasoconstrictionVasodilators (ACEi, ARB, hydralazine)
Contractility (inotropy)Intrinsic force at same preloadCatecholamines, digoxin, Ca2+, dobutamineBeta-blockers, heart failure, acidosis, hypoxia

Frank-Starling Law

The Frank-Starling law states that stroke volume increases with increased preload (end-diastolic volume), up to a point. Greater stretch of cardiac muscle fibers → greater force of contraction due to optimal actin-myosin overlap.

⭐ Boards ApplicationIn heart failure, the Frank-Starling curve is shifted DOWN and to the RIGHT — same preload produces less stroke volume. Inotropes shift the curve UP (more SV at same preload). This explains why HF patients decompensate with fluid challenges.
🧐 Quick Formula ReviewCO = HR x SV. MAP = DBP + 1/3 (SBP-DBP). SVR = (MAP - RAP) / CO x 80. Ejection fraction = SV/EDV (normal >55%).

Blood Pressure and Hemodynamics

BP = CO x SVR (systemic vascular resistance). Arterioles are the primary site of resistance. The baroreceptor reflex (carotid sinus, aortic arch) provides moment-to-moment BP regulation.

Pressure TypeNormal ValueClinical Note
Systolic BP90-120 mmHgReflects CO and arterial compliance
Diastolic BP60-80 mmHgReflects SVR and arterial tone
MAP70-100 mmHgPerfusion pressure; organs autoregulate within this range
Pulse pressureSBP - DBP (~40 mmHg)Widened in aortic regurgitation, aortic dissection; narrowed in cardiac tamponade, severe AS
CVP (RAP)2-8 mmHgReflects preload; elevated in RHF, tamponade, tension pneumo

Autonomic Regulation of the Heart

EffectSympathetic (NE, Epi)Parasympathetic (ACh)
Heart rateIncrease (beta-1)Decrease (M2)
ContractilityIncrease (beta-1)Slight decrease (atria mainly)
AV conductionFaster (shorter PR)Slower (longer PR; AV block with excess)
Coronary vesselsDilate (beta-2); constrict (alpha-1)Dilate (indirect via metabolic demand)
🩹 Valsalva Maneuver PhasesStrain phase: increased intrathoracic pressure → decreased venous return → decreased CO → reflex tachycardia. Release phase: venous return surges → CO spike → reflex bradycardia. Used to terminate SVT (increases vagal tone).

Clinical Pearls

⭐ Ejection Fraction ClassificationHFrEF: EF less than 40% (systolic dysfunction). HFmrEF: EF 40-49%. HFpEF: EF 50% or greater (diastolic dysfunction). HFpEF is now more common than HFrEF; associated with hypertension, obesity, DM, elderly women.
🩹 Starling Forces (Capillary Exchange)Filtration favored by: high capillary hydrostatic pressure (Pc), low interstitial oncotic pressure. Absorption favored by: high plasma oncotic pressure (albumin), low Pc. Edema: when filtration exceeds lymphatic drainage.
🧐 Orthostatic Hypotension ReflexStanding: gravity pools blood in legs → decreased venous return → decreased CO → baroreceptors detect BP drop → sympathetic activation → tachycardia + vasoconstriction → BP restored. Failure = orthostatic hypotension (20 mmHg systolic or 10 mmHg diastolic drop).