Physiology

Gallbladder Physiology

Bile storage, concentration, CCK-mediated secretion, and enterohepatic circulation

Bile Composition and Function

Bile is produced by hepatocytes (~500-1000 mL/day) and stored/concentrated in the gallbladder (10x concentration via water/electrolyte absorption). Primary function: emulsification of dietary fats to aid lipase digestion.

ComponentSourceFunction
Bile saltsHepatocytes (from cholesterol)Emulsify fats; form micelles for fat-soluble vitamin absorption (A,D,E,K)
Bilirubin (conjugated)Hepatocytes (from heme)Waste product; gives stool brown color
CholesterolHepatocytesExcreted in bile; excess → gallstones
Phospholipids (lecithin)HepatocytesSolubilize cholesterol in bile; prevent gallstones
Water and electrolytesBile ductulesVehicle; adjusted by secretin and aldosterone

CCK and Secretin Regulation

HormoneStimulusSourceGallbladder Effect
CCK (cholecystokinin)Fat and protein in duodenumI cells (duodenum)Gallbladder contraction + sphincter of Oddi relaxation → bile release
SecretinAcid (H+) in duodenumS cells (duodenum)Stimulates bile duct bicarbonate secretion (watery bile); augments CCK effect
SomatostatinProtein, fat, acidD cells (pancreas, gut)Inhibits CCK, gastrin, secretin; decreases bile secretion
⭐ Boards PearlGallbladder contraction requires CCK. Acalculous cholecystitis risk: TPN use (no CCK stimulus → bile stasis → infection). Vagotomy also impairs gallbladder emptying.

Enterohepatic Circulation

Bile salts are reabsorbed in the terminal ileum (95% actively reabsorbed; 5% lost in stool) → portal vein → liver → re-secreted into bile. Total pool circulates 6-8x per day.

🩹 Clinical RelevanceTerminal ileum resection (Crohn's disease) → bile salt malabsorption → steatorrhea (fat malabsorption) + bile salt diarrhea (cholerheic diarrhea). Cholestyramine binds bile salts → used for cholestatic itch and hypercholesterolemia.
🧐 Gallstone TypesCholesterol stones (80%): supersaturated bile + nucleation; risk factors: female, fat, fertile, forty, fair (5 Fs). Pigment stones: black (hemolysis — sickle cell, hereditary spherocytosis); brown (infection — Clonorchis, E. coli).

Clinical Pearls

⭐ Murphy's SignInspiratory arrest on deep palpation of right upper quadrant = Murphy's sign = acute cholecystitis. Sonographic Murphy's sign (tenderness with ultrasound probe) is highly specific. Choledocholithiasis: stone in CBD → obstructive jaundice + elevated ALP and direct bilirubin + acholic stools + dark urine.
⭐ Charcot's Triad vs Reynold's PentadCharcot's triad (ascending cholangitis): RUQ pain + fever/chills + jaundice. Reynold's pentad: + altered mental status + hypotension (septic shock — emergency ERCP needed).